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writing exist than his description of the coronation procession | of Anne Boleyn through the streets of London, few more full of picturesque power than that in which he relates how the spire of St Paul's was struck by lightning; and to have once read is to remember for ever the touching and stately words in which he compares the monks of the London Charterhouse preparing for death with the Spartans at Thermopylae. Proofs of his power in the sustained narration of stirring events are abundant; his treatment of the Pilgrimage of Grace, of the sea fight at St Helens and the repulse of the French invasion, and of the murder of Rizzio, are among the most conspicuous examples of it. Nor is he less successful when recording pathetic events, for his stories of certain martyrdoms, and of the execution of Mary queen of Scots, are told with exquisite feeling and in language of well-restrained emotion. And his characters are alive. We may not always agree with his portraiture, but the men and women whom he saw exist for us instinct with the life with which he endows them and animated by the motives which he attributes to them. His successes must be set against his failures. At the least he wrote a great history, one which can never be disregarded by future writers on his period, be their opinions what they may; which attracts and delights a multitude of readers, and is a splendid example of literary form and grace in historical composition.

The merits of his work met with full recognition. Each instalment of his History, in common with almost everything which he wrote, was widely read, and in spite of some adverse criticisms was received with eager applause. In 1868 he was elected rector of St Andrews University, defeating Disraeli by a majority of fourteen. He was warmly welcomed in the United States, which he visited in 1872, but the lectures on Ireland which he delivered there caused much dissatisfaction. On the death of his adversary Freeman in 1892, he was appointed, on the recommendation of Lord Salisbury, to succeed him as regius professor of modern history at Oxford. Except to a few Oxford men, who considered that historical scholarship should have been held to be a necessary qualification for the office, his appointment gave general satisfaction. His lectures on Erasmus and other 16th-century subjects were largely attended. With some allowance for the purpose for which they were originally written, they present much the same characteristics as his earlier historical books. His health gave way in the summer of 1894, and he died on the 20th of October.

His long life was full of literary work. Besides his labours as an author, he was for fourteen years editor of Fraser's Magazine. He was one of Carlyle's literary executors, and brought some sharp criticism upon himself by publishing Carlyle's Reminiscences and the Memorials of Jane Welsh Carlyle, for they exhibited the domestic life and character of his old friend in an unpleasant light. Carlyle had given the manuscripts to him, telling him that he might publish them if he thought it well to do so, and at the close of his life agreed to their publication. Froude therefore declared that in giving them to the world he was carrying out his friend's wish by enabling him to make a posthumous confession of his faults. Besides publishing these manuscripts he wrote a Life of Carlyle. His earlier study of Irish history afforded him suggestions for a historical novel entitled The Two Chiefs of Dunboy (1889). In spite of one or two stirring scenes it is a tedious book, and its personages are little more than machines for the enunciation of the author's opinions and sentiments. Though Froude had some intimate friends he was generally reserved. When he cared to please, his manners and conversation were charming. Those who knew him well formed a high estimate of his ability in practical affairs. In 1874 Lord Carnarvon, then colonial secretary, sent Froude to South Africa to report on the best means of promoting a confederation of its colonies and states, and in 1875 he was again sent to the Cape as a member of a proposed conference to further confederation. Froude's speeches in South Africa were rather injudicious, and his mission was a failure (see SOUTH AFRICA: History). He was twice married. His first wife, a

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daughter of Pascoe Grenfell and sister of Mrs Charles Kingsley, died in 1860; his second, a daughter of John Warre, M.P. for Taunton, died in 1874. Froude's Life, by Herbert Paul, was published in 1905.

(W. Hu.) FRUCTOSE, LAEVULOSE, or FRUIT-SUGAR, a carbohydrate of the formula C6H12O6. It is closely related to ordinary dglucose, with which it occurs in many fruits, starches and also in honey. It is a hydrolytic product of inulin, from which it may be prepared; but it is more usual to obtain it from "invert sugar," the mixture obtained by hydrolysing cane sugar with sulphuric acid. Cane sugar then yields a syrupy mixture of glucose and fructose, which, having been freed from the acid and concentrated, is mixed with water, cooled in ice and calcium hydroxide added. The fructose is precipitated as a saccharate, which is filtered, suspended in water and decomposed by carbon dioxide. The liquid is filtered, the filtrate concentrated, and the syrup so obtained washed with cold alcohol. On cooling the fructose separates. It may be obtained as a syrup, as fine, silky needles, a white crystalline powder, or as a granular crystalline, somewhat hygroscopic mass. When anhydrous it melts at about 95° C. It is readily soluble in water and in dilute alcohol, but insoluble in absolute alcohol. It is sweeter than cane sugar and is more easily assimilated. It has been employed under the name diabetin as a sweetening agent for diabetics, since it does not increase the sugar-content of the urine, other medicinal applications are in phthisis (mixed with quassia or other bitter), and for children suffering from tuberculosis or scrofula in place of cane sugar or milk-sugar.

Chemically, fructose is an oxyketone or ketose, its structural formula being CH2OH (CH-OH)3 CO-CH2OH; this result followed from its conversion by H. Kiliani into methylbutylacetic acid. The form described above is laevo-rotatory, but it is termed d-fructose, since it is related to d-glucose. Solutions exhibit mutarotation, fresh solutions having a specific rotation of 104.0°, which gradually diminishes to -92°. It was synthesized by Emil Fischer, who found the synthetic sugar which he named a-acrose to be (d+1)-fructose, and by splitting this mixture he obtained both the d and forms. Fructose resembles d-glucose in being fermentable by yeast (it is the one ketose which exhibits this property), and also in its power of reducing alkaline copper and silver solutions; this latter property is assigned to the readiness with which hydroxyl and ketone groups in close proximity suffer oxidation. For the structural (stereochemical) relations of fructose see SUGAR.

FRUGONI, CARLO INNOCENZIO MARIA (1692-1768), Italian poet, was born at Genoa on the 21st of November 1692. He was originally destined for the church and at the age of fifteen, in opposition to his strong wishes, was shut up in a convent; but although in the following year he was induced to pronounce monastic vows, he had no liking for this life. He acquired considerable reputation as an elegant writer both of Latin and Italian prose and verse; and from 1716 to 1724 he filled the chairs of rhetoric at Brescia, Rome, Genoa, Bologna and Modena successively, attracting by his brilliant fluency a large number of students at each university. Through Cardinal Bentivoglio he was recommended to Antonio Farnese, duke of Parma, who appointed him his poet laureate; and he remained at the court of Parma until the death of Antonio, after which he returned to Genoa. Shortly afterwards, through the intercession of Bentivoglio, he obtained from the pope the remission of his monastic vows, and ultimately succeeded in recovering a portion of his paternal inheritance. After the peace of Aix-laChapelle he returned to the court of Parma, and there devoted the later years of his life chiefly to poetical composition. He died on the 20th of December 1768. As a poet Frugoni was one of the best of the school of the Arcadian Academy, and his lyrics and pastorals had great facility and elegance.

His collected works were published at Parma in 10 vols. in 1799. and a more complete edition appeared at Lucca in the same year in 15 vols. A selection from his works was published at Brescia in 1782, in 4 vols.

FRUIT (through the French from the Lat. fructus; frui, to enjoy), in its widest sense, any product of the soil that can be enjoyed by man or animals; the word is so used constantly in the Bible, and extended, as a Hebraism, to offspring or progeny of man and of animals, in such expressions as "the fruit of the body,' "" of the womb,” ,""fruit of thy cattle " (Deut. xxviii. 4), &c., and generally to the product of any action or effort. Between this wide and frequently figurative use of the word and its application in the strict botanical sense treated below, there is a popular meaning, regarding the objects denoted by the word entirely from the standpoint of edibility, and differentiating them roughly from those other products of the soil, which, regarded similarly, are known as vegetables. In this sense "fruit" is applied to such seed-envelopes of plants as are edible, either raw or cooked, and are usually sweet, juicy or of a refreshing flavour. But applications of the word in this sense are apt to be loose and shifting according to the fashion of the time.

Fruit, in the botanical sense, is developed from the flower as the result of fertilization of the ovule. After fertilization various changes take place in the parts of the flower. Those more immediately concerned in the process, the anther and stigma, rapidly wither and decay, while the filaments and style often remain for some time; the floral envelopes become dry, the petals fall, and the sepals are either deciduous, or remain persistent in an altered form; the ovary becomes enlarged, forming the pericarp; and the ovules are developed as the seeds, containing the embryo-plant. The term fruit is strictly applied to the mature pistil or ovary, with the seeds in its interior; but it often includes other parts of the flower, such as the bracts and floral envelopes. Thus the fruit of the hazel and oak consists of the ovary enveloped by the bracts; that of the apple and pear, of the ovary and floral receptacle; and that of the pineapple, of the whole inflorescence. Such fruits are sometimes distinguished as pseudocarps. In popular language, the fruit includes all those parts which exhibit a striking change as the result of| fertilization. In general, the fruit is not ripened unless fertilization has been effected; but cases occur as the result of cultivation in which the fruit swells and becomes to all appearance perfect, while no seeds are produced. Thus, there are seedless oranges, grapes and pineapples. When the ovules are unfertilized, it is common to find that the ovary withers and does not come to maturity; but in the case of bananas, plantains and breadfruit, the non development of seeds seems to lead to a larger growth and a greater succulence of fruit.

its formation. In the gooseberry (fig. 29), grape, guava, tomato and pomegranate, the seeds nestle in pulp formed by the placentas. In the orange the pulpy matter surrounding the seeds is formed by succulent cells, which are produced from the inner partitioned lining of the pericarp. In the strawberry the receptacle becomes succulent, and bears the mature carpels on its convex surface (fig. 2): in the rose there is a fleshy hollow receptacle which bears the carpels on its concave surface (fig. 3). In the juniper the scaly bracts grow up round the seeds and become succulent, and in the fig (fig. 4) the receptacle becomes succulent and encloses an inflorescence.

The fruit, like the ovary, may be formed of a single carpel or of several. It may have one cell or cavity, being unilocular; or many, multilocular, &c. The number and nature of the divisions depend on the number of carpels and the extent to which their edges are folded inwards. The appearances presented by the ovary do not always remain permanent in the fruit. Great changes are observed to take place, not merely as regards the increased size of the ovary, its softening or hardening, but also in its internal structure, owing to the suppression, additional formation or enlargement of parts. Thus, in the ash (fig. 1) an ovary with two cells, each containing an ovule attached to a central placenta, is changed into a unilocular fruit with one seed; one ovule becomes abortive, while the other, g, gradually enlarging until the septum is pushed to one side, unites with the walls of the cell, and the placenta appears to be parietal. In the oak and hazel, an ovary with three and two cells respectively, and two ovules in each, produces a one-celled fruit with one seed. In the coco-nut, a trilocular and triovular ovary produces a onecelled, one-seeded fruit. This abortion may depend on the pressure caused by the development of certain ovules, or it may proceed from non-fertilization of all the ovules and consequent non-enlargement of the carpels. Again, by the growth of the placenta, or the folding inwards of parts of the carpels, divisions occur in the fruit which did not exist in the ovary. In Cathartocarpus Fistula a one-celled ovary is changed into a fruit having each of its seeds in a separate cell, in consequence of spurious dissepiments being produced horizontal from the inner wall of the ovary. In flax (Linum) by the folding inwards of the back of the carpels a five-celled ovary becomes a ten-celled fruit. In Astragalus the folding inwards of the dorsal suture converts a one-celled ovary into a two-celled fruit; and in Oxytropis the folding of the ventral suture gives rise to a similar change. The development of cellular or pulpy matter, and the enlargement of parts not forming whorls of the flower, frequently alter the appearance of the fruit, and render it difficult to discover

The pericarp consists usually of three layers, the external, or epicarp (fig. 5, ep); the middle, or mesocarp, m; and the internal,

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FIG. 1.-Samara or winged fruit of Ash (Fraxinus). 1, Entire, with its wing a; 2, lower portion cut transversely, to show that it consists of two cells; one of which, l, is abortive, and is reduced to a very small cavity, while the other is much enlarged and filled with a seed g. FIG. 2.-Fruit of the Strawberry (Fragaria vesca), consisting of an enlarged succulent receptacle, bearing on its surface the small dry seed-like fruits (achenes). (After Duchartre.)

From Strasburger's Lehrbuch der Botanik, by permission of Gustav Fischer.

FIG. 3.-Fruit of the Rose cut vertically. s', Fleshy hollowed receptacle; s, persistent sepals; fr, ripe carpels; e, stamens, withered,

FIG. 4-Peduncle of Fig (Ficus Carica), ending in a hollow receptacle enclosing numerous male and female flowers.

FIG. 5-Fruit of Cherry (Prunus Cerasus) in longitudinal section. ep, Epicarp; m, mesocarp; en, endocarp.

From Strasburger's Lehrbuch der Botanik, by permission of Gustav Fischer. or endocarp, en. These layers are well seen in such a fruit as the peach, plum or cherry, where they are separable one from the other; in them the epicarp forms what is commonly called the skin; the mesocarp, much developed, forms the flesh or pulp, and hence has sometimes been called sarcocarp; while the endocarp, hardened by the production of woody cells, forms the stone or putamen immediately covering the kernel or seed. The pulpy matter found in the interior of fruits, such as the gooseberry, grape and others, is formed from the placentas, and must not be confounded with the sarcocarp. In some fruits, as in the nut, the three layers become blended together and are indistinguishable. In bladder senna (Colutea arborescens) the pericarp retains its leaflike appearance, but in most cases it becomes altered both in consistence and in colour. Thus in the date the epicarp is the outer brownish skin, the pulpy matter is the mesocarp or sarcocarp, and the thin papery-like lining is the endocarp covering the hard seed. In the medlar the endocarp becomes of a stony hardness. In the melon the epicarp and endocarp are very thin, while the mesocarp forms the bulk of the fruit, differing in texture and taste in its external and internal parts. The rind of the orange consists of epicarp and mesocarp, while the endocarp forms partitions in the interior, filled with pulpy cells. The part of the pericarp attached to the peduncle is the base, and the point where the style or stigma existed is the apex. This latter is not always the apparent apex, as in the case of the ovary; it may be lateral or even basilar. The style sometimes remains in a hardened form, rendering the fruit apiculate; at other times it falls off, leaving only traces of its existence. The presence of the style or stigma serves to distinguish certain single. seeded pericarps from seeds.

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united carpels, two types of dehiscence occur a longitudinal and a transverse. In the longitudinal the separation may take place by the dissepiments throughout their length, so that the fruit is resolved into its original carpels, and each valve represents a carpel, as in rhododendron, Colchicum, &c.; this dehiscence, in consequence of taking place through the septum, is called septicidal (figs. 9, 10). The valves separate from their commissure, or central line of union, carrying the placentas with them, or they leave the latter in the centre, so as to form with the axis a column of a cylindrical, conical or prismatic shape. Dehiscence is loculicidal when the union between the edges of the carpels is persistent, and they dehisce by the dorsal suture, or through the back of the loculaments, as in the lily and iris (figs. 11, 12). In these cases each valve consists of a half of each of two contiguous carpels. The placentas either remain united to the axis, or they separate from it, being attached to the septa on the valves. When the outer walls of the carpels break off from the septa, leaving them attached to the central column, the dehiscence is said to be septifragal (fig. 13), and where, as in Linum catharticum and Calluna, the splitting takes place first of all in a septicidal manner, the fruit is described as septicidally septifragal; while in other cases, as in thorn apple (Datura Stramonium), where the splitting is at first loculicidal, the dehiscence is loculicidally septifragal. In all those forms the separation of the valves takes place either from above downwards or from below upwards. In itni anabose bonitos to stiul lo song rive coal deindaide Adenogailo yndlu da ni gibis betion whose or basablen

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The pod stub (legume) of the Pea; 7, the dorsal suture; b, the ventral; c, calyx; s, seeds. Vanl From Vines' Students' Text-Book of Botany, by perqsaaste mission of Swan Sonnenschein & Co.

elaqen F10. 9. (1) Fruit or capsule of Meadow lons Saffron (Colchicum autumnale), dehiscing along vide no the septa (septicidally); (2) same cut across, showing the three chambers with the seeds nattached along the middle line (axile placenmol adal zonetation).cam niemen ffene FIG. 11. FIG. 10.-Diagram to illustrate the septiviss bombom) cidal dehiscence in a pentalocular capsule. The loculaments / correspond to the number of the carpels, which separate by splitting through the septa, s.id BFIG. 11.-The seed vessel (capsule) of the Flower-de-Luce (Iris), opening in a loculicidal manner. The three valves bear the septa in the centre, and the opening takes place through the back of the loculaments. Each valve is formed by the halves of contiguous carpels, and todos os

FIG. 12.-Diagram to illustrate loculicidal dehiscence. The loculaments, split at the back, and the valves separate, bearing the septa s on their centres. lose it iw boy

FIG. 13.-Diagram to illustrate septifragal dehiscence, in which the dehiscence takes place through the back of the loculaments 1, and the valves separate from the septa s, which are left attached to the placentas in the centre.

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seen in the order Euphorbiaceae, where there are usually three such carpels, and the fruit is termed tricoccus. In many of them, as Hura crepitans, the cocci separate with great force and elasticity. In many leguminous plants, such as Ornithopus, Hedysarum (fig. 17), Entada, Coronilla and the gum-arabic plant (Acacia arabica), the fruit becomes a schizocarp by the formation of transverse partitions from the folding in of the sides of the pericarp, and distinct separations taking place at these partitions. Fruits are formed by one flower, or are the product of several flowers combined. In the former case they are either apocarpous, of one mature carpel or of several separate free carpels: or syncarpous, of several carpels, more or less completely united. When the fruit is composed of the ovaries of several flowers united, it is usual to find the bracts and floral envelopes also joined with them, so as to form one mass; hence such fruits are known as multiple, confluent or anthocarpous. The term simple is applied to fruits which are formed by the ovary of a single flower, whether they are composed of one or several carpels, and whether these carpels are separate or combined.

seed.

The object of the fruit in the economy of the plant is the protection and nursing of the developing seed and the dispersion of the ripe seeds. Hence, generally, one-seeded fruits are indehiscent, Dispersal while fruits containing more than one seed open to allow of fruit or of the dispersal of the seeds over as wide an area as possible. The form, colour, structure and method of dehiscence of fruits and the form of the contained seeds are intimately associated with the means of dispersal, which fall into several categories. (1) By a mechanism residing in the fruit. Thus many fruits open suddenly when they are dry, and the seeds are ejected by the twisting or curving of the valves, or in some other way; e.g. in gorse, by the spiral curving of the valves; in Impatiens, by the twisting of the cocci; in squirting cucumber, by the pressure exerted on the pulpy contents by the walls of the pericarp. (2) By aid of various external agencies such as water. Fruits or seeds are sometimes sufficiently buoyant to float for a long time on seaor fresh-water; e.g. coco-nut, by means of its thick, fibrous coat (mesocarp), is carried hundreds of miles in the sca, the tough, leathery outer coat (epicarp) preventing it from becoming watersoaked. Fruits and seeds of West Indian plants are thrown up on the coasts of north-west Europe, having been carried by the Gulf Stream, and will often germinate; many are rendered buoyant by air-containing cavities, and the embryo is protected from the seawater by the tough coat of fruit or seed. Water-lily seeds are surrounded with a spongy tissue when set free from the fruit, and float for some distance before dropping to the bottom. (3) The most general agent in the dispersal of seeds is the wind or currents of air-the fruit or seed being rendered buoyant by wing-developments as in fruits of ash (fig. 1) or maple (fig. 21), seeds of pines and firs, or many members of the order Bignoniaceae; or hairdevelopinents as in fruits of clematis, where the style forms a feathery

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FIG. 19.

FIG. 20.

FIG. 21.

From Vines' Students' Text-Book of Botany, by permission of Swan Sonnenschein & Co.

FIG. 19.-Dry one-seeded fruit of dock (Rumex) cut vertically. ov, Pericarp formed from ovary wall; s, seed; e, endosperm; pl, embryo with radicle pointing upwards and cotyledons downwardsenlarged.

FIG. 20.-Achene of Ranunculus arvensis in longitudinal section; e, endosperm; pl, embryo. (After Baillon, enlarged.)

From Strasburger's Lehrbuch der Botanik, by permission of Gustav Fischer. FIG. 21-Fruit of Common Sycamore (Acer Pseudoplatanus), dividing into two mericarps m; s, pedicel; fl, wings (nat. size). appendage, fruits of many Compositae (dandelion, thistle, &c.). which are crowned by a plumose pappus, or seeds of willow and poplar, or Asclepias (fig. 36), which bear tufts of silky hairs; to this category belong bladder-like fruits, such as bladder-senna, which are easily rolled by the wind, or cases like the so-called rose

of Jericho, a small cruciferous plant (Anastatica hierocuntica), where the plant dries up after developing its fruits and becomes detached from the ground; the branches curl inwards, and the whole plant is rolled over the dry ground by the wind. The wind also aids the dispersal of the seeds in the case of fruits which open by small teeth (many Caryophyllaceae [fig. 6]) or pores (poppy [fig. 7), Campanula. &c.); the seeds are in these cases small and numerous, and are jerked through the pores when the capsules, which are generally borne on long, dry stems or stalks, are shaken by the wind. (4) In other cases members of the animal world aid in seed-dispersal. Fruits often bear stiff hairs or small hooks, which cling to the coat of an animal or the feathers of a bird; such are fruits of cleavers (Galium A parine), a common hedge-row plant, Ranunculus arvensis (fig. 20), carrot, Geum, &c.; or the fruit or seed has an often bright-coloured, fleshy

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FIG. 22.-Vertical section of a grain of wheat, showing embryo below at the base of the endosperm e; s, scutellum separating embryo from endosperm; fl, foliage leaf; p.s, sheath of plumule; p.r, primary root; s.p.r, sheath of primary root.

calyx, c. The style s appears to arise from the base of the carpels, FIG. 23. Fruit of Comfrey (Symphytum) surrounded by persistent enlarged.

FIG. 24. Ovary of Foeniculum officinale with pendulous ovules, in longitudinal section. (After Berg and Schmidt, magnified.)

From Strasburger's Lehrbuch der Botanik, by permission of Gustav Fischer. fruit. The two carpels have separated so as to form two mericarps FIG. 25.-Fruit of Carum Carui. A, Ovary of the flower; B, ripe (m). Part of the septum constitutes the carpophore (a). p, Top of flower-stalk; d, disk on top of ovary; n, stigma.

"

Forms of fruit.

From Vines' Students' Text-Book of Botany, by permission of Swan Sonnenschein & Co. covering, which is sought by birds as food, as in stone-fruits such as plum, cherry (fig. 5), &c., where the seed is protected from injury in the mouth or stomach of the animal by the hard endocarp; or the hips of the rose (fig. 3), where the succulent scarlet "fruit (the swollen receptacle) envelops a number of small dry true fruits (achenes), which cling by means of stiff hairs to the beak of the bird. Simple fruits have either a dry or succulent pericarp. The achene is a dry, one-seeded, indehiscent fruit, the pericarp of which is closely applied to the seed, but separable from it. It is solitary, forming a single fruit, as in the dock (fig. 19) and in the cashew, where it is supported on a fleshy peduncle; or aggregate, as in Ranunculus (fig. 20), where several achenes are placed on a common elevated receptacle. In the strawberry the achenes (fig. 2) are aggregated on a convex succulent receptacle. In the rose they are supported on a concave receptacle (fig. 3), and in the fig the succulent receptacle completely encloses the achenes (fig. 4). In Dorstenia the achenes are situated on a flat or slightly concave receptacle. Hence what in common language are called the seeds of the strawberry, rose and fig, are in reality ripe carpels. The styles occasionally remain attached to the achenes in the form of feathery appendages, as in Clematis. In Compositae, the fruit is an inferior achene (cypsela), to which the pappus (modified calyx) remains adherent. Such is also the nature of the fruit in Dipsacaceae (e.g. scabious). When the pericarp is thin, and appears like a bladder surrounding the seed, the achene is termed a utricle, as in Amarantaceae. When the pericarp is extended in the form of a winged appendage, a samara or samaroid achene is produced, as in the ash (fig. 1) and common sycamore (fig. 21). In these cases there are usually two achenes united, one of which, however, as in Fraxinus (fig. 1), may be abortive. The wing sur rounds the fruit longitudinally in the clm. When the pericarp be comes so incorporated with the seed as to be inseparable from it. as in grains of wheat (fig. 22), maize, oats and other grasses, then the name caryopsis is given. The one-seeded 'portions (mericarps) of schizocarps often take the form of achenes, e.g. the mericarps of the

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bellso einela odini mallows or of umbellifers (figs. 24, 25). In Labiatae and Boraginaceae (e.g. comfrey, fig. 23), where the bicarpellary ovary becomes our one-seeded portions in the fruit, the partial fruits are of the nature of achenes or nutlets according to the texture (leathery or hard) of the pericarp.

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-gl

The nut or glans is a dry one-celled indehiscent fruit with a hardened pericarp, often surrounded by bracts at the base, and, when mature, containing only one seed. In the young state the ovary often contains two or more ovules, but only one comes to maturity. It is illustrated by the fruits of the hazel and chestnut, which are covered by leafy bracts, in the form of a husk, and by the acorn, in which the bracts and receptacle form a cupula or cup (fig. 26). The parts of the pericarp of the nut are united so as to appear one. In common language the term nut is very vaguely applied both to fruit and seeds, The drupe is a succulent usually one-seeded indehiscent fruit, with a pericarp easily distinguishable into epicarp, mesocarp and endocarp. This term is applied to such fruits as the cherry (fig. 5), peach, plum, apricot or mango. The endocarp is usually hard, forming the stone (putamen) of the fruit, which encloses the kernel or seed. The mesocarp is generally pulpy and succulent, so as to be truly a sarcocarp, as in the peach, but it is sometimes of a tough texture, as in the almond, and at other times is more or less fibrous, as in the coco-nut. In the almond there are often two ovules formed, only one of which comes to perfection. In the raspberry and bramble several small drupes or drupels are aggregated so as to

From Strasburger's Lehrbuch der Botanik, by permission of Gustav Fischer. FIG. 26.-Cupule of Quercus Aegilops. cp, cupule; gl, fruit. (After Duchartre.)

constitute an elaerio.

The follicle is a dry unilocular many-seeded fruit, formed from one carpel and dehiscing by the ventral suture. It is rare to meet with a solitary follicle forming the fruit. There are usually several aggregated together, either in a whorl on a shortened receptacle, as in hellebore, aconite, larkspur, columbine (figs. 27, 28) or the order Crassulaceae, or in a spiral manner on an elongated receptacle, as in Magnolia and Banksia. Occasionally, follicles dehisce by the dorsal suture, as in Magnolia grandiflora and Banksia.

The legume or pod is a dry monocarpellary unilocular many-seeded fruit, formed from one carpel, dehiscing both by the ventral and the

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The berry (bacca) is a term applied generally to all fruits with seeds immersed in pulp, and includes fruits of very various origin. In Actaea (baneberry) or Berberis duo iu oda har (barberry) it is derived from ar single free carpel; generally, how day ever, it is the product of a syn-d bu carpous ovary, which is superior, yo as in grape or potato, or inferior, as in gooseberry (fig. 29) or currant. In the pomegranate there is a peculiar baccate many-celled inferior fruit, having a tough rind, enclosing two rows of carpels placed one above the other. The seeds are immersed in pulp, and are attached irregularly to the wall, base and centre of the loculi. In the baobab there is a multi

FIG. 31.-Transverse section locular syncarpous fruit, in which of the fruit of the Melon the seeds are immersed in pulp. (Cucumis Melo), showing the The pepo, another indehiscent placentas with the seeds attached syncarpous fruit, is illustrated by to them. The three carpels the fruit of the gourd, melon (fig. forming the pepo are separated 31) and other Cucurbitaceae. It by partitions. From the centre is formed of three carpels, sur- processes pass outwards, ending mounted by the calyx; the rind in the curved placenta. is thick and fleshy, and there are three or more sced-bearing parietal placentas, either surrounding a central cavity or prolonged inwards into it. The fruit of the papaw resembles the pepo, but the calyx is not superior. The hesperidium is the name given to such indehiscent fleshy syncarpous fruits as the orange, lemon and shaddock, in which the epicarp and mesocarp form a separable rind, and the endocarp sends prolongations inwards, forming triangular divisions, to the inner angle of which the seeds are attached, pulpy cells being developed around them from the wall. Both pepo and hesperidium may be considered as modifications of the berry.

The pome (fig. 30), seen in the apple, pear, quince, medlar and hawthorn, is a fleshy indehiscent syncarpous fruit, in the formation of which the receptacle takes part. The outer succulent part is the swollen receptacle, the horny core being the true fruit developed from the usually five carpels and enclosing the seeds. In the medlar the core (or true pericarp) is of a stony hardness, while the outer succulent covering is open at the summit. The pome somewhat resembles the fruit of the rose (fig. 3), where the succulent receptacle surrounds a number of separate achenes.

The name capsule is applied generally to all dry syncarpous fruits, which dehisce by valves. It may thus be unilocular or multilocular, one or many-seeded. The true valvular capsule is observed in Colchicum (fig. 9), lily and iris (fig. 11). The porose capsule is seen in the poppy (fig. 7), Antirrhinum and Campanula. In Campanula the pores occur at the base of the capsule, which becomes inverted when ripe. When the capsule opens by a lid, or by circum scissile 350 dehiscence, it is called a pyxidium, as in pimpernel (A nagallis atom arvensis) (fig, 16), henbane and monkey-pot (Lecythis). The capsule assumes a screw-like form in Helicteres, and a star-like form in starou anise (Illicium anisatum). In certain instances the cells of the Sue capsule separate from each other, and open with elasticity to scatter me the seeds. This kind of capsule is met with in the sandbox tree Bevel (Hura crepitans) and other Euphorbiaceae, where the cocci, conertaining each a single seed, burst asunder with force; and in Geraniaceae, where the cocci, each containing, when mature, usually one seed, separate from the carpophore, become curved upwards by their adherent styles, and open by the ventral suture (fig. 18).

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FIG. 27.-Fruit of Columbine (Aquilegia), formed of five follicles: FIG. 28.-Single follicle, showing dehiscence by the ventral suture. FIG. 29.-Transverse section of berry of Gooseberry, showing the seeds attached to the parietal placentas and immersed in pulp, which is formed partly from the endocarp, partly from the seed-coat. FIG. 30.-Section of the fruit of the Apple (Pyrus Malus), or pome, consisting of a fleshy covering formed by the floral receptacle and the true fruit or core with five cavities with seeds.

dorsal suture. It characterizes leguminous plants, as the bean and pea (fig. 8). In the bladder-senna it forms an inflated legume. In some Leguminosae, as Arachis, Cathartocarpus Fistula and the tamarind, the fruit must be considered a legume, although it does not dehisce. The first of these plants produces its fruit underground, and is called earth-nut; the second has a partitioned legume and is schizocarpic; and both the second and third have pulpy matter surrounding the seeds. Some legumes are schizocarpic by the formation of constrictions externally. Such a form is the lomentum or lomentaceous legume of Hedysarum (fig. 17), Coronilla, Ornithopus, Entada and of some Acacias. In Medicago the legume

from two carpels, with a false septum, dehiscing by two valves The siliqua is a dry syncarpous bilocular many-seeded fruit, formed from below upwards, the valves separating from the placentas and leaving them united by the septum (fig. 32). The seeds are attached on both sides of the septum, either in one row or in two. When the fruit is long and narrow it is a siliqua (fig. 14); when broad and short, silicula (fig. 33). It occurs in cruciferous plants, as wallflower, cabbage and cress. In Glaucium and Eschscholtzia (Papaveraceae) the dissepiment is of a spongy nature. It may become transversely constricted (lomentaceous), as in radish (Raphanus) and sea-kale, and it may be reduced, as in woad (Isatis), to a oneseeded condition.

It sometimes happens that the ovaries of two flowers unite so as to form a double fruit (syncarp). This may be seen in many species of honeysuckle. But the fruits which are now to be considered consist usually of the floral envelopes, as well as the ovaries of several flowers united into one, and are called multiple or confluent. The term anthocarpous has also been applied as indicating that the floral envelopes as well as the carpels are concerned in the formation of the fruit.

The sorosis is a succulent multiple fruit formed by the confluence

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