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positions, figures, colours. The prospect from Hampstead-hill is compressed into the compass of a six-pence, yet circumstantially represented. A stage coach, travelling at its ordinary speed for half an hour, passes, in the eye, only over one twelfth of an inch, yet is this change of place in the image distinctly perceived throughout its whole progress; for it is only by means of that perception that the motion of the coach itself is made sensible to the eye. If any thing can abate our admiration of the smallness of the visual tablet compared with the extent of vision, it is a reflection which the view of nature leads us, every hour, to make, viz. that, in the hands of the Creator, great and little are nothing.

Sturmius held, that the examination of the eye was a cure for atheism. Besides that conformity to optical principles which its internal constitution displays, and which alone amounts to a manifestation of intelligence having been exerted in the structure; besides this, which forms, no doubt, the leading character of the organ, there is to be seen, in every thing belonging to it and about it, an extraordinary degree of care, an anxiety for its preservation, due, if we may so speak, to its value and its tenderness. It is lodged in a strong, deep, bony socket, composed by the junction of seven different bones*, hollowed out at their edges. In some few species, as that of the coatimondi t, the orbit is not bony throughout; but when ever this is the case, the upper, which is the deficient part, is supplied by a cartilaginous ligament; a substitution which shows the same care. Within this socket it is imbedded in fat, of all animal substances the best adapted both to its repose and motion. It is sheltered by the eyebrows; an arch of hair, which, like a thatched penthouse, prevents the sweat and moisture of the forehead from running down into it.

But it is still better protected by its lid. Of the superficial parts of the animal frame, I know none which, in its office and structure, is more deserving of attention than the eyelid. It defends the eye; it wipes it; it closes it in sleep.

Are

there, in any work of art whatever, purposes more evident than those which this organ fulfils? or an apparatus for executing those purposes more intelligible,

* Heister, sect. 89.

more appropriate, or more mechanical? If it be overlooked by the observer of nature, it can only be because it is obvious and familiar. This is a tendency to be guarded against. We pass by the plainest instances, whilst we are exploring those which are rare and curious; by which conduct of the understanding, we sometimes neglect the strongest observations, being taken up with others, which, though more recondite and scientific, are, as solid arguments, entitled to much less consideration.

In order to keep the eye moist and clean (which qualities are necessary to its brightness and its use), a wash is constantly supplied by a secretion for the pur pose; and the superfluous brine is conveyed to the nose through a perforation in the bone as large as a goose-quill. When once the fluid has entered the nose, it spreads itself upon the inside of the nostril, and is evaporated by the current of warm air, which, in the course of respiration, is continually passing over it. Can any pipe or outlet, for carrying off the waste liquor from a dye-house or a distillery, be more mechanical than this is? It is easily perceived, that the eye must want moisture; but could the want of the eye generate the gland which produces the tear, or bore the hole by which it is discharged,—a hole through a bone?

It is observable, that this provision is not found in fish,—the element in which they live supplying a constant lotion to the eye.

It were, however, injustice to dismiss the eye as a piece of inechanism, without noticing that most exquisite of all contrivances, the nictitating membrane, which is

found in the eyes of birds and of many quadrupeds. Its use is to sweep the eye, which it does in an instant, to spread over it the lachrymal humour; to defend it also from sudden injuries; yet not totally, when drawn upon the pupil, to shut out the light.

The commodiousness with which it lies folded up in the upper corner of the eye, ready for use and action, and the quickness with which it executes its purpose, are properties known and obvious to every observer; but what is equally admirable, though not quite so obvious, is the combination of two kinds of substance, muscular and elastic, and of two different kinds of action, by which the

+ Mem. R. Ac. Paris, p. 117,

motion of this membrane is performed. It is not, as in ordinary cases, by the action of two antagonist muscles, one pulling forward and the other backward, that a reciprocal change is effected; but it is thus: The membrane itself is an elastic substance, capable of being drawn out by force like a piece of elastic gum, and by its own elasticity returning, when the force is removed, to its former position. Such being its nature, in order to fit it up for its office, it is connected by a tendon or thread with a muscle in the back part of the eye: this tendon or thread, though strong, is so fine, as not to obstruct the sight, even when it passes across it; and the muscle itself, being placed in the back part of the eye, derives from its situation the advantage, not only of being secure, but of being out of the way; which it would hardly have been in any position that could be assigned to it in the anterior part of the orb, where its function lies. When the muscle behind the eye contracts, the membrane, by means of the communicating thread, is instantly drawn over the fore-part of it. When the muscular contraction (which is a positive, and, most probably, a voluntary effort) ceases to be exerted, the elasticity alone of the membrane brings it back again to its position. Does not this, if any thing can do it, bespeak an artist, master of his work, acquainted with his materials? "Of a thousand other things," say the French Academicians, we perceive not the contrivance, because we understand them only by the effects, of which we know not the causes: but we here treat of a machine, all the parts whereof are visible; and which need only be looked upon, to discover the reasons of its motion and action t."

66

In the configuration of the muscle which, though placed behind the eye, draws the nictitating membrane over the eye, there is, what the authors, just now quoted, deservedly call a marvellous mechanism. I suppose this structure to be found in other animals; but, in the memoirs from which this account is taken, it is anatomically demonstrated only in the cassowary. The muscle is passed through a loop formed by another muscle; and is there inflected, as if it were round a

* Philos. Transact. 1796.

1

pulley. This is a peculiarity; and observe the advantage of it. A single muscle with a straight tendon, which is the common muscular form, would have been sufficient, if it had had power to draw far enough. But the contraction, necessary to draw the membrane over the whole eye, required a longer muscle than could lie straight at the bottom of the eye. Therefore, in order to have a greater length in a less compass, the cord of the main muscle makes an angle. This, so far, answers the end; but, still farther, it makes an angle, not round a fixed pivot, but round a loop formed by another muscle; which second muscle, whenever it contracts, of course twitches the first muscle at the point of inflection, and thereby assists the action designed by both.

One question may possibly have dwelt in the reader's mind during the perusal of these observations, namely, Why should not the Deity have given to the animal the faculty of vision at once? Why this circuitous perception; the ministry of so many means; an element provided for the purpose; reflected from opaque substances, refracted through transparent ones; and both according to precise laws; then, a complex organ, an intricate and artificial apparatus, in order, by the operation of this element, and in conformity with the restrictions of these laws, to produce an image upon a membrane communica ting with the brain? Wherefore all this? Why make the difficulty in order to surmount it? If to perceive objects by some other mode than that of touch, or objects which lay out of the reach of that sense, were the thing proposed; could not a simple volition of the Creator have communicated the capacity? Why resort to contrivance, where power is omnipotent? Contrivance, by its very definition and nature, is the refuge of imperfection. To have recourse to expedients, implies difficulty, impediment, restraint, defect of power. This question belongs to the other senses, as well as to the sight; to the general functions of animal life, as nutrition, secretion, respiration; to the œconomy of vegetables; and indeed to almost all the operations of nature. The question, therefore, is of very wide extent; and amongst other answers which

Memoirs for a Natural History of Animals, by the Royal Academy of Sciences at Paris, done into English by Order of the Royal Society, 1701, page 249.

and

I

may be given to it, besides reasons of which probably we are ignorant, one answer is this: It is only by the display of contrivance, that the existence, the agency, the wisdom of the Deity, could be testiThis is the fied to his rational creatures. scale by which we ascend to all the knowledge of our Creator which we possess, so far as it depends upon the phænomena, or the works of nature. Take away this, from us every subject you take away of observation, and ground of reasoning; mean as our rational faculties are formed at present. Whatever is done, God could have done without the intervention of instruments or means: but it is in the construction of instruments, in the choice and adaptation of means, that a creative intelligence is seen. It is this which constitutes the order and beauty of the universe. God, therefore, has been pleased to prescribe limits to his own power, and to work his ends within those limits. The general laws of matter have perhaps the nature of these limits; its inertia, its reaction; the laws which govern the communication of motion, the refraction and reflection of light, the constitution of fluids, non-elastic and elastic, the transmission of sound through the latter; the laws of magnetism, of electricity; and probably others, yet undiscovered. These are general laws; and when a particular purpose is to be effected, it is not by making a new law, nor by the suspension of the old ones, nor by making them wind, and bend, and yield to the occasion (for nature with great steadiness adheres to and supports them); but it is, as we have seen in the eye, by the interposition of an apparatus, corresponding with these laws, and suited to the exigency which results from them, that the purpose is at length attained.

As we

as

ranks of these. We do not advance this
a doctrine either of philosophy or of
religion; but we say that the subject may
safely be represented under this view;
because the Deity, acting himself by
general laws, will have the same conse-
quences upon our reasoning, as if he had
prescribed these laws to another. It has
been said, that the problem of creation
was, "attraction and matter being given,
to make a world out of them :" and, as
above explained, this statement perhaps
does not convey a false idea.

have said, therefore, God prescribes limits
to his power, that he may let in the exer-
cise, and thereby exhibit demonstrations,
of his wisdom. For then, i. e. such
laws and limitations being laid down, it is
as though one Being should have fixed
certain rules; and, if we may so speak,
provided certain materials; and, after
wards, have committed to another Being,
out of these materials, and in subordina-
tion to these rules, the task of drawing
forth a creation: a supposition which evi-
dently leaves room, and induces indeed a
necessity for contrivance. Nay, there
may be many such agents, and many

eye

We have made choice of the eye as an instance upon which to rest the argument offered of this chapter. Some single example was to be proposed: and the itself under the advantage of admitting of a strict comparison with optical instruThe ear, it is probable, is no ments. less artificially and mechanically adapted to its office than the eye. But we know less about it: we do not so well understand the action, the use, or the mutual dependency of its internal parts. Its general form, however, both external and internal, is sufficient to show that it is an instrument adapted to the reception of sound; that is to say, already knowing that sound consists in pulses of the air, we perceive, in the structure of the ear, a suitableness to receive impressions from this species of action, and to propagate these impressions to the brain. For of what does this structure consist? An external ear (the concha), calculated, like an ear-trumpet, to catch and collect the pulses of which we have spoken; in large quadrupeds, turning to the sound, and possessing a configuration, as well as motion, evidently fitted for the office: of a tube which leads into the head, lying at the root of this outward ear, the folds and sinuses thereof tending and conducting the air towards it; of a thin membrane, like the pelt of a drum, stretched across this passage upon a bony rim: of a chain of moveable, and infinitely curious bones, forming a communication, and the only communication, that can be observed, between the membrane last mentioned and the interior channels and recesses of the skull: of cavities, similar in shape and form to wind instruments of music, being spiral or portions of circles: of the eustachian tube, like the hole in a drum, to let the air pass freely into and out of the barrel of the ear, as the covering membrane vibrates, or as the temperature may

be altered; the whole labyrinth hewn out of a rock; that is, wrought into the substance of the hardest bone of the body. This assemblage of connected parts constitutes together an apparatus, plainly enough relative to the transmission of sound, or of the impulses received from sound, and only to be lamented in not being better understood.

diminishing the space through which it oscillates; both of which changes may augment or facilitate the still deeper action of the auditory nerves.

The benefit of the eustachian tube to the organ may be made out upon known pneumatic principles. Behind the drum of the ear is a second cavity, or barrel, called the tympanum. The eustachian The communication within, formed by tube is a slender pipe, but sufficient for the small bones of the ear, is, to look the passage of air, leading from this upon, more like what we are accustomed cavity into the back part of the mouth. to call machinery, than any thing I am Now, it would not have done to have had acquainted with in animal bodies. It It a vacuum in this cavity; for, in that case, seems evidently designed to continue to- the pressure of the atmosphere from withwards the sensorium the tremulous mo- out would have burst the membrane which tions which are excited in the membrane covered it. Nor would it have done to of the tympanum, or what is better known have filled the cavity with lymph or any by the name of the "drum of the ear." other secretion; which would necessarily The compages of bones consists of four, have obstructed, both the vibration of the which are so disposed and so hinge up- membrane, and the play of the small on one another, as that if the membrane, bones. Nor, lastly, would it have done the drum of the ear, vibrate, all the four to have occupied the space with confined are put in motion together; and, by the air, because the expansion of that air by result of their action, work the base of heat, or its contraction by cold, would that which is the last in the series, upon have distended or relaxed the covering an aperture which it closes, and upon membrane, in a degree inconsistent with which it plays, and which aperture opens the purpose which it was assigned to execute. The only remaining expedient, and that for which the eustachian tube serves, is to open to this cavity a communication with the external air. In one word; it exactly answers the purpose of the hole in a drum.

into the tortuous canals that lead to the brain. This last bone of the four is called the stapes. The office of the drum of the ear is to spread out an extended surface, capable of receiving the impressions of sound, and of being put by them into a state of vibration. The office of the stapes is to repeat these vibrations. It is a repeating frigate, stationed more within the line. From which account of its action may be understood, how the sensation of sound will be excited, by any thing which communicates a vibratory motion to the stapes, though not, as in all ordinary cases, through the intervention of the membrana tympani. This is done by solid bodies applied to the bones of the skull, as by a metal bar holden at one end between the teeth, and touching at the other end a tremulous body. It likewise appears to be done, in a considerable degree, by the air itself, even when this membrane, the drum of the ear, is greatly damaged. Either in the natural or preternatural state of the organ, the use of the chain of bones is to propagate the impulse in a direction towards the brain, and to propagate it with the advantage of a lever: which advantage consists in increasing the force and strength of the vibration, and at the same time

The membrana tympani itself likewise, deserves all the examination which can be made of it. It is not found in the ears of fish; which furnishes an additional proof of what indeed is indicated by every thing about it, that it is appropriated to the action of air, or of an elastic medium. It bears an obvious resemblance to the pelt or head of a drum, from which it takes its name. It resembles also a drum-head in this principal property, that its use depends upon its tension. Tension is the state essential to it. Now we know that, in a drum, the pelt is carried over a hoop, and braced as occasion requires, by the means of strings attached to its circumference. In the membrane of the ear, the same purpose is

provided for, more simply, but not less mechanically, nor less successfully, by a different expedient, viz. by the end of a bone (the handle of the malleus) pressing upon its centre. It is only in very large animals that the texture of this membrane can be discerned.

In the.

Philosophical Transactions for the year 1800 (vol. i.), Mr. Everard Home has given some curious observations upon the ear, and the drum of the ear of an elephant. He discovered in it, what he calls a radiated muscle, that is, straight muscular fibres, passing along the membrane from the circumference to the centre; from the bony rim which surrounds it towards the handle of the malleus to which the central part is attached. This muscle he supposes to be designed to bring the membrane into unison with different sounds: but then he also discovered, that this muscle itself cannot act, unless the membrane be drawn to a stretch, and kept in a due state of tightness, by what may be called a foreign force, viz. the action of the muscles of the malleus. Supposing his explanation of the use of the parts to be just, our author is well founded in the reflection which he makes upon it: "that this mode of adapting the ear to different sounds, is one of the most beautiful applications of muscles in the body; the mechanism is so simple, and the variety of effects so great."

In another volume of the Transactions above referred to, and of the same year, two most curious cases are related of persons who retained the sense of hearing, not in a perfect, but in a very considerable degree, notwithstanding the almost total loss of the membrane we have been describing. In one of these cases, the use here assigned to that membrane, of modifying the impressions of sound by change of tension, was attempted to be supplied by straining the muscles of the outward ear. "The external ear," we are told, "had acquired a distinct motion upward and backward, which was observable whenever the patient listened to any thing which he did not distinctly hear: when he was addressed in a whisper, the ear was seen immediately to move: when the tone of voice was louder, it then remained altogether motionless."

It appears probable, from both these cases, that a collateral, if not principal, use of the membrane, is to cover and protect the barrel of the ear which lies behind it. Both the patients suffered from cold: one, "a great increase of deafness from catching cold;" the other, "very considerable pain from exposure to a stream of cold air." Bad effects therefore followed from this cavity being left open to the external air; yet, had the Author of nature shut it up by

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The generation of the animal no more accounts for the contrivance of the eye or ear, than, upon the supposition stated in a preceding chapter, the production of a watch by the motion and mechanism of a former watch, would account for the skill and attention evidenced in the watch, so produced; than it would account for the disposition of the wheels, the catching of their teeth, the relation of the several parts of the works to one another, and to their common end, for the suitableness of their forms and places to their offices, for their connexion, their operation, and the useful result of that operation. I do insist most strenuously upon the correctness of this comparison; that it holds as to every mode of specific propagation; and that whatever was true of the watch, under the hypothesis above mentioned, is true of plants and animals.

1. To begin with the fructification of plants. Can it be doubted but that the seed contains a particular organization? Whether a latent plantule with the means of temporary nutrition, or whatever else it be, it encloses an organization suited to the germination of a new plant. Has the plant which produced the seed any thing more to do with that organization, than the watch would have had to do with the structure of the watch which was produced in the course of its mechanical movement? I mean, Has it any thing at all to do with the contrivance? The maker and contriver of one watch, when he inserted within it a mechanism suited to the production of another watch, was, in truth, the maker and contriver of that other watch. All the properties of the new watch were to be referred to his agency: the design manifested in it, to his intention: the art, to him as the artist: the collocation of each part, to his placing: the action, effect, and use, to his counsel, intelligence, and workmanship. In producing it by the intervention of a former watch, he was only working by one set of tools instead of another. So it is with the plant, and the seed produced by it. Can any

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